Optimus Prime was the last born of the original Thirteen Transformers. His spark was the most like their progenitor Primus, its frequency nearly identical. He was the mediator and visionary of the group, passing among them all as a calm, inspiring, thoughtful, unifying and well-liked friend. It was his unique spark and his inspiring reassurance that "All are One" that allowed the Primes to rally and succeed in their battle against the Chaos Bringer Unicron. When tragedy at last ended the era of the Primes and brought forth the new race of lesser descendant Transformers he alone chose to be reborn in the Well of All Sparks as one of them, that he might know them and their needs more completely.
All memory of his past life gone, he took the name Orion Pax and sought his way like any other robot on the new world. Eventually he found his place as a simple data clerk, satisfied with his job. He was always anxious of the inequality amongst Cybertronians and thus, after gaining inspiration from an outspoken gladiator named Megatron's call for an upheaval of Cybertron's society, took a stand to rid the oppressive caste system which hindered the freedom of all the sentient beings on Cybertron. Little did he realize that his instructor, the master archivist Alpha Trion, was also one of the Thirteen and knew of the secret within the data clerk. Once Pax and Megatron discovered they did not have as much in common as they once believed, the lowly data clerk was promoted by the High Council to be the next Prime. With the claiming of the Matrix of Leadership Orion Pax became Optimus Prime once more, regaining his memory and rising to lead his people.
Over the centuries, loss and war have led Optimus to appear stern and serious to most. But scratch the surface, and you'll find an individual who greatly cares for his fellow living beings, with a warmth and kindness his closest friends can rely on. You'll also find regret over the loss of a comrade to darkness, and the faintest glimmer of hope that he can be saved.
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